On the Origin of Species

On the Origin of Species

Title page of the 1859 edition[1]
Author	Charles Darwin
Country	United Kingdom of Great Britain and Ireland
Language	English
Subject	Natural selection Evolutionary biology
Published	24 November 1859[2] (John Murray)
Media type	Print (hardback)
Pages	502
OCLC	352242
Preceded by	On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection
Followed by	Fertilisation of Orchids
Text	On the Origin of Species at Wikisource

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Evolutionary biology
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Index Introduction Main Outline Glossary Evidence History
Processes and outcomes Population genetics Variation Diversity Mutation Natural selection Adaptation Polymorphism Genetic drift Gene flow Speciation Adaptive radiation Co-operation Coevolution Coextinction Divergence Convergence Parallel evolution Extinction
Natural history Origin of life Common descent History of life Timeline of evolution Human evolution Phylogeny Biodiversity Biogeography Classification Evolutionary taxonomy Cladistics Transitional fossil Extinction event
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On the Origin of Species (or, more completely, On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life)^[3] is a work of scientific literature by Charles Darwin that is considered to be the foundation of evolutionary biology; it was published on 24 November 1859.^[4] Darwin's book introduced the scientific theory that populations evolve over the course of generations through a process of natural selection although Lamarckism was also included as a mechanism of lesser importance. The book presented a body of evidence that the diversity of life arose by common descent through a branching pattern of evolution. Darwin included evidence that he had collected on the Beagle expedition in the 1830s and his subsequent findings from research, correspondence, and experimentation.^[5] The opening sentences demonstrate that Darwin did not see the role of natural selection in species conformity or what is now called stabilizing selection. This was not corrected in editions subsequent to Darwin reading the correct explanation In Patrick Matthew's book in 1860 which had been published in 1831.

Various evolutionary ideas had already been proposed to explain new findings in biology. There was growing support for such ideas among dissident anatomists and the general public, but during the first half of the 19th century the English scientific establishment was closely tied to the Church of England, while science was part of natural theology. Ideas about the transmutation of species were controversial as they conflicted with the beliefs that species were unchanging parts of a designed hierarchy and that humans were unique, unrelated to other animals. The political and theological implications were intensely debated, but transmutation was not accepted by the scientific mainstream.

The book was written for non-specialist readers and attracted widespread interest upon its publication. Darwin was already highly regarded as a scientist, so his findings were taken seriously and the evidence he presented generated scientific, philosophical, and religious discussion. The debate over the book contributed to the campaign by T. H. Huxley and his fellow members of the X Club to secularise science by promoting scientific naturalism. Within two decades, there was widespread scientific agreement that evolution, with a branching pattern of common descent, had occurred, but scientists were slow to give natural selection the significance that Darwin thought appropriate. During "the eclipse of Darwinism" from the 1880s to the 1930s, various other mechanisms of evolution were given more credit. With the development of the modern evolutionary synthesis in the 1930s and 1940s, Darwin's concept of evolutionary adaptation through natural selection became central to modern evolutionary theory, and it has now become the unifying concept of the life sciences.